In review or revision
LM Hennelly, Shivam Shrotriya, S Khan, A Mohammadi, M Farhadinia, L Llaneza, J Bump, A Homkes, S Windels, MZ Islam, YV Jhala, JV Lopez-Bao, G Roffler, M Godbole, I Ghorpade, P Comizzoli, N Songsasen, H Sand, C Wikenros, P Wabakken, B Zimmermann, P Mahoney, D Stahler, E Stahler, M Metz, K Cassidy, T Gable, B Habib . In review. Population-specific drivers of reproductive phenology in a widespread carnivore, the gray wolf. Ecosphere (https://www.biorxiv.org/content/biorxiv/early/2026/05/26/2026.05.25.727694.full.pdf)
Sampedro A, LM Hennelly, A Koshkina, M Girtsina, N Marmazinskaya, S Li, S Kachel, A Nawaz, M Chetri, C Auguliaro, B Nasanbat, S Lyngdoh, A Mukhashov, T Dendup, V Terentyev, J Robinson, J Janecka, A Barashkova, F Rovero, H Malvandi, I Cancellare, H Hojamuradov, T Kholikov, JV Lopez-Bao, G Werhahn. In review. Wolves in Central Asia: Elevation delineates boundaries across wolf lineages. Biological Conservation
NG Martins, X Sun, LT Lanigan , LM Hennelly, CF Alemany, JR Madrigal, I Svetilcic, A Galov, C Fernandes, HE Alqamy, SO Ozkurt, Z Krikova, G Goldthorpe, G Gachal, SM Madjdzadeh, C Nowak, M Niedzialkowska, P Hulva, L Paule, R Myslajek, S Nowak, U Saarma, J Ozolins, M Murtskhvaladze, F Suchentrunk, D Murariu, M Krofel, M Heltai, E Geffen, BN Sacks, W Bogdanowicz, MHS Sinding, MTP Gilbert, S Gopalakrishnan. In revision. Unraveling the golden jackal’s genomic journey: insights into their origin, expansion, and hybridization across Eurasia from whole-genome data. Iscience
Todd ET, Fontsere C, Sun X, Scharff-Olsen CH, Hernández-Alonso G, Lanigan LT, Martins NFG, Ciucani MM, Ramos-Madrigal J, Hennelly L, Mak SST, Andersone-Lilley Z, Åsberg A, Balčiauskas L, Baltrūnaitė L, Baryshnikov GF, Boldbaatar B, Boldgiv B, Černá Bolfíková B, Borowik T, Bujnáková D, Ciucci P, Çoban AK, Çoban E, Erlandsson M, Flagstad Ø, Frantz L, Geffen E, Harmoinen J, Jelk L, Kalthoff DC, Karamanlidis AA, Kemahlı-Aytekin MÇ, Kojola I, Kopatz A, Kosintsev P, Kusak J, Kuznetsova A, Kvist L, Larsson L, Larson G, Linderholm A, Lingaas F, Llaneza L, Martin SLF, Meiri S, Øverli Ø, Paule L, Sablin M, Skage M, Stålfjäll C, Stanton DWG, Şekercioğlu ÇH, Špinkyte R, Tirronen K, Törngren L, Vernesi C, Yamaguchi N, Åkesson M, Mysłajek RW, Hulva P, Nowak C, Niedziałkowska M, Nowak S, Dalén L, Godinho R, Saarma U, Fumagalli L, Aspi J, Stenøien HK, Martin MD, Sinding MHS, Gilbert MTP, Gopalakrishnan S. 2026. The population structure and genetic health of European wolves. (https://www.biorxiv.org/content/10.64898/2026.03.20.712003v1.full.pdf).
B Habib, S Khan, S Sadhukhan, G Sarwar, S Yellapu, S Viranta, LM Hennelly. submitted. Population estimation of Indian wolves (Canis lupus pallipes) in the Indian subcontinent. Oryx
Published
20. Pecnerova P, Ishida Y, Garcia-Erill G, Bertola LD, Santander CG, Liu X, Bruniche-Olsen A, Khan A, Hennelly LM, Balboa RF, Lin L, Rasmussen MS, Wang X, Schubert M, Al-Chaer A, Urbaniak S, Nobuta K, Treadup S, Viaud-Martinez KA, Flamingh A, Malhi RS, Bird S, Ting N, Matsa M, Tchamba MN, Bourgeois S, Thouless C, Douglas-Hamilton I, Wittemyer G, Demeke Y, Groot PHVC, Muwanika VB, Masembe C, Nyakaana S, Georgiadis NJ, Hvilson C, Heller R, Siegismund JR, Roca AL. 2026. The genomic impact of connectivity and decline in Africa’s elephants. Nature Communications
19. Hennelly LM, MHS Sinding. Dogs have deep genetic roots in ice-age Europe. 2026. Nature
18. Hennelly LM, Barbara R Parreira, Ash Noble, Camilla Scharff-Olsen, M. Çisel Kemahlı Aytekin, Çağan H. Şekercioğlu, Pavel Kosintsev, Ladislav Paule, Pavel Hulva, Hans Stenoien, Bilal Habib, Hira Fatima, Ghulam Sarwar, Frank Hailer, Xin Sun, Nuno Filipes Gomes Martin, M Thomas P Gilbert, Benjamin N Sacks, Mikkel-Holger S. Sinding, Shyam Gopalakrishnan. 2025. Continent-wide view of genomic diversity and divergence in the wolves of Asia. Communications Biology
17. Battilani D, JR Madrigal, LM Hennelly, S Gopalakrishnan, C Vernesi, F Mattucci, E Fabbri, P Ciucci, R Caniglia. 2025. Whole-genome data to investigate recent and historical dog introgression patterns in Italian wolves. Ecology and Evolution. 15(12): e72508
16. Parreira BR, P Pecnerova, L Tensen, Aninta SG, Bruniche-Olsen A, Khan A, Morales HE, LM Hennelly. 2025. Q&A: Inbreeding and its implications for conservation. BMC Biology
15. G Werhahn, C Auguliaro, M Kabir, LM Hennelly, M Chetri, HA Hikmani, A Mohammadi, YV Jhala, DW Macdonald, MS Farhadinia. 2025. Asia’s wolves and synergies with big cats. Conservation Letters
14. Cancellare IA, B Weckworth, A Caragiulo, KL Pilgrom, TM McCarthy, A Abdullaev, A Abdullaev, X Bian, E Bykova, I Dais-Freedman, M Gritsina, LM Hennelly, S Janjua, OJ Johansson, S Kachel, A Karnaukhov, M Korablev, Z Kubanychbekov, R Kulenbekov, X Liang, P Lkhagvajav, TK Meyer, B Munkhtsog, B Munkhtsog, MA Nawaz, S Ostrowski, M Paltsyn, A Poyarkov, S Rabinowitz, T Rooney, T Rosen, VV Rozhnov, BN Sacks, MK Schwartz, KP McCarthy. 2025. Pleistocene refugia and the impacts on genetic variability in High Asia: range-wide phylogeography of snow leopards. Biodiversity and Conservation
13. LM Hennelly, G Sarwar, H Fatima, G Werhahn, S Katchel, FI Abbas, AM Khan, Z Kubanychbekov, T Mahmood, MR Khan, H Aisha, M Waseem, M Farooq, BN Sacks. 2024. Genomic analysis of wolves from Pakistan clarifies boundaries among three divergent wolf lineages. Journal of Heredity esad066
12. H Fatima, T Mahmood, LM Hennelly, M Farooq, N Munawar, W Ali, BN Sacks. 2023. Distributional patterns and habitat association of sympatric carnivores in Margalla Hills National Park, Pakistan, and a comparison of conventional versus identification in carnivore scatology. Animal Biology 1:1-20
11. Plassais J, BM vonHoldt, HG Parker, A Carmagnini, N Dubos, I Papa, K Bevant, T Derrien, LM Hennelly, DT Whitaker, AC Harris, AN Hogan, HJ Huson, VF Zaibert, A Linderholm, J Haile, T Fest, B Habib, BN Sacks, N Benecke, AK Outram, MV Sablin, M Germonpre, G Larson, L Frantz, EA Ostrander. 2022. Natural and human-driven selection of a single non-coding body size variant in ancient and modern canids. Current Biology 32(4): 889-897
10. LM Hennelly, B Habib, S Modi, EK Rueness, P Gaubert, BN Sacks. 2021. Ancient divergence of Indian and Tibetan wolves revealed by recombination-aware phylogenomics. Molecular Ecology 30:6687-6700
9. Rafaqat I, T Mahmood, LM Hennelly, F Akrim, IU Zaman, H Fatima, S Andleeb, G Muhammad, Y Abbas, M Farooq, N Munawar, MR Khan, A Hamid,MA Orakzai. 2021. Phylogenetic status and diet of red fox (Vulpes vulpes griffithii) inhabiting Ayubia National Park, Pakistan. Brazilian Journal of Biology 84:e252991.
8. Sarwar G, AM Khan, F Abbas, MT Wasim, LM Hennelly. 2021. First record on body morphometrics and chemical immobilization of wolves from Pakistan. Pakistan Veterinary Journal http://dx.doi.org/10.29261/pakvetj/2021.054
7. Sacks BN, KJ Mitchell, CB Quinn, LM Hennelly, MHS Sinding, MJ Statham, S Preckler-Quisquater, SR Fain, L Kistler, SL Vanderzwan, JA Meachen, EA Ostrander, LAF Frantz. 2021. Pleistocene origins, western ghost lineages, and the emerging phylogeographic history of the red wolf and coyote. Molecular Ecology 30(17):4292-4304
6. Fatima H, T Mahmood, LM Hennelly, M Farooq, F Akrim, MS Nadeem. 2021. Spatial distribution and dietary niche breadth of leopard cat (Prionailurus bengalensis) inhabiting Margalla Hills National Park, Pakistan. Zoological Studies 60:34
5. Weyna A, RR Dubielzig, LM Hennelly. 2020. The relative importance of phylogeny and habitat in determining the presence and prominence of a granula iridica in hooved mammals. Veterinary Ophthalmology 1-8.
4. Sadhukhan S, LM Hennelly, B Habib. 2019. Characterising the vocal repertoire of the Indian wolf (Canis lupus pallipes). PloS ONE 14(10): e021618
3. Hamid A, T Mahmood, H Fatima, LM Hennelly, R Akrim, A Hussain, M Waseem. 2019. Origin, ecology, and human conflict of grey wolf (Canis lupus) in Suleiman Range, South Waziristan, Pakistan. Mammalia 83(6): 539-551.
2. Hennelly LM, B Habib, H Root-Gutteridge, V Palacios, D Passilongo. 2017. Howl variation across Himalayan, North African, Indian and Holarctic wolf clades: tracing divergence in the world’s oldest wolf lineages using acoustics. Current Zoology 63(3): 341-348
1. Hennelly LM, B Habib, and S Lyngdoh. 2015. Himalayan wolf and feral dog displaying mating behavior from Spiti Valley, India and potential conservation threats from sympatric feral dogs. Canid Biology & Conservation 18(9): 33-36.
Hennelly Lab 